It turns out that Greenland sharks routinely live to be 272 years old. One was caught recently that may have been 512 years old. That is to say, she would have been born in 1504.
She had been swimming in the northern sea, starting only 12 years after Columbus discovered America. She would have been 13 years old when Martin Luther posted his theses. She could have eaten a Pilgrim.
This would make this species of shark the longest-living vertebrate. Scientists are trying to figure out how these creatures can live so long, hoping to apply their findings to human beings.
UPDATE: The shark, whose long life was ignominiously ended when it was caught in a fishing net, was female. So I have changed the earlier pronoun “he” to “she.” Also, as the linked story says, the scientists determined that the shark was between 272 and 512 years old, probably more likely 400 than the upper limit. But still, that’s old.